• The recipient of much praise and acclaim, Veterinary Parasitology is widely considered to be the definitive veterinary parasitology reference for practitioners and students alike. This Fourth Edition has been developed and enhanced into a two-part reference to reflect recent advances in the field, modern teaching practice, and updated parasite taxonomic classification systems.

    Part One contains expanded individual parasite descriptions using current taxonomic status within three new chapters on Helminthology, Protozoology and Entomology. Further updated chapters are provided on: The laboratory diagnosis of parasitism, Antiparasitics, The epidemiology of parasitic diseases, and Host resistance to parasitic diseases. Host species chapters have been retained and expanded and are found in Part Two of the edition.

    KEY FEATURES

    • Tailored for those directly involved in the diagnosis, treatment and control of parasitic diseases of domestic animals
    • Compatible with the diversity of current parasitology teaching modules – both for teaching parasite systematics and diseases on a host-organ basis
    • Offers the most detailed parasite descriptions available today for teachers, research groups, veterinarians in practice and in government service, and others involved in aspects of parasitic disease

    Thoroughly revised and restructured to reflect the most up-to-date advancements in the field, Veterinary Parasitology, Fourth Edition, enhances its stellar reputation as the gold standard reference text for the global veterinary profession.

  • Preface to the first edition, xxi

    Acknowledgements to the first edition, xxii

    Preface and acknowledgements to the second edition, xxiii

    Preface and acknowledgements to the third edition, xxiv

    Preface and acknowledgements to the fourth edition, xxv

    (Contents list continues on pages vii–xx)

    Part 1: General parasitology including taxonomy, diagnosis, antiparasitics

    1 VETERINARY HELMINTHOLOGY

    2 VETERINARY PROTOZOOLOGY

    3 VETERINARY ENTOMOLOGY

    4 Laboratory diagnosis of parasitism, 259

    Helminth infections, 259

    Faecal examination, 259

    Collection of faeces, 259

    Methods of examination of faeces, 259

    Identification of nematode eggs, 261

    Identification of trematode eggs, 261

    Larval recovery, 268

    Recovery of alimentary nematodes, 269

    Key to the identification of gastrointestinal nematodes of ruminants, 271

    Recovery of adult lungworms, 274

    Recovery of trematode and cestode parasites, 275

    Other aids to diagnosis of ruminant nematodes, 275

    DNA]based methods, 276

    Protozoal infections, 283

    Examination of faecal samples for coccidia, 283

    Examination of faecal samples for other protozoa, 283

    Examination of blood and lymph, 289

    Examination of skin, 290

    Xenodiagnosis, 290

    Ectoparasites, 290

    Insects, 290

    Ticks and mites, 301

    Mounting and preservation, 309

    Chemicals and solutions, 311

    Flotation solutions, 311

    Worm count solutions, 311

    Stains, 311

    5 Antiparasitics, 313

    Anthelmintics, 313

    Anthelmintics and their mode of action, 313

    Properties of anthelmintic compounds, 317

    Use of anthelmintics, 317

    Methods of administration, 318

    Antiprotozoals, 318

    Antiprotozoals and their mode of action, 319

    Use of antiprotozoals, 323

    Methods of administration, 323

    Ectoparasiticides (insecticides/acaricides), 323

    Ectoparasiticides and their mode of action, 324

    Methods of pesticide application and uses, 327

    Parasiticide resistance, 328

    Anthelmintic resistance, 328

    Antiprotozoal resistance, 333

    Pesticide resistance, 334

    6 The epidemiology of parasitic diseases, 336

    An increase in the numbers of infective stages, 336

    Contamination of the environment, 336

    Development and survival of infective stages, 337

    An alteration in host susceptibility, 338

    Altered effects of an existing infection, 338

    Altered susceptibility to the acquisition of new infections, 340

    The introduction of susceptible stock, 340

    Absence of acquired immunity, 340

    Absence of age immunity, 341

    Longevity of infective stages, 341

    Influence of genetic factors, 341

    Strain of parasite, 341

    Introduction of infection into a clean environment, 341

    Introduction of new stock, 341

    Role of effluent, 341

    Role of infected vectors, 342

    7 Host resistance to parasitic diseases, 343

    Species resistance, 343

    Age resistance, 343

    Breed resistance, 343

    Acquired immunity to helminth infections, 344

    Effect of the immune response, 344

    Evasion of the host’s immune response, 345

    Cost of the immune response, 346

    Acquired immunity to protozoal infections, 346

    Acquired immunity to arthropod infections, 347

    The future of parasite vaccines, 347

    Helminths, 347

    Protozoa, 349

    Ectoparasites, 351

    Part 2: Host–parasite diseases

    8 Parasites of cattle, 352

    Endoparasites, 352

    Parasites of the digestive system, 352

    Parasites of the respiratory system, 379

    Parasites of the liver, 383

    Parasites of the pancreas, 388

    Parasites of the circulatory system, 389

    Parasites of the nervous system, 409

    Parasites of the reproductive/urogenital system, 410

    Parasites of the locomotory system, 413

    Parasites of the connective tissue, 417

    Parasites of the integument, 420

    Ectoparasites, 424

    Flies, 424

    Lice, 424

    Mites, 427

    Cattle parasite checklist, 431

    9 Parasites of sheep and goats, 436

    Endoparasites, 436

    Parasites of the digestive system, 436

    Parasites of the respiratory system, 474

    Parasites of the liver, 480

    Parasites of the pancreas, 489

    Parasites of the circulatory system, 489

    Parasites of the nervous system, 497

    Parasites of the reproductive/urogenital system, 498

    Parasites of the locomotory system, 500

    Parasites of the integument, 502

    Ectoparasites, 503

    Lice, 503

    Mites, 506

    Fly strike (myiasis), 511

    Sheep parasite checklist, 516

    Goat parasite checklist, 520

    10 Parasites of horses, 524

    Endoparasites, 524

    Parasites of the digestive system, 524

    Parasites of the respiratory system, 544

    Parasites of the liver, 545

    Parasites of the circulatory system, 545

    Parasites of the nervous system, 552

    Parasites of the reproductive/urogenital system, 554

    Parasites of the locomotory system, 555

    Parasites of the integument, 556

    Ectoparasites, 558

    Lice, 558

    Mites, 559

    Horse parasite checklist, 561

    11 Parasites of pigs, 565

    Endoparasites, 565

    Parasites of the digestive system, 565

    Parasites of the respiratory system, 583

    Parasites of the liver, 584

    Parasites of the pancreas, 585

    Parasites of the circulatory system, 586

    Parasites of the nervous system, 588

    Parasites of the reproductive/urogenital system, 588

    Parasites of the locomotory system, 590

    Parasites of the integument, 592

    Ectoparasites, 592

    Lice, 592

    Mites, 593

    Pig parasite checklist, 596

    12 Parasites of dogs and cats, 599

    Endoparasites, 599

    Parasites of the digestive system, 599

    Parasites of the respiratory system, 629

    Parasites of the liver, 635

    Parasites of the circulatory system, 640

    Parasites of the nervous system, 654

    Parasites of the reproductive/urogenital system, 656

    Parasites of the locomotory system, 657

    Parasites of the integument, 659

    Ectoparasites, 662

    Lice, 662

    Mites, 663

    Dog parasite checklist, 670

    Cat parasite checklist, 674

    13 Parasites of poultry and gamebirds, 678

    Endoparasites, 678

    Parasites of the digestive system, 678

    Parasites of the respiratory system, 720

    Parasites of the liver, 723

    Parasites of the circulatory system, 723

    Parasites of the nervous system, 729

    Parasites of the reproductive/urogenital system, 729

    Parasites of the locomotory system, 731

    Parasites of the integument, 732

    Ectoparasites, 733

    Lice, 733

    Mites, 737

    Chicken parasite checklist, 742

    Turkey parasite checklist, 746

    Duck parasite checklist, 749

    Goose parasite checklist, 752

    Pheasant parasite checklist, 754

    Partridge parasite checklist, 756

    Quail parasite checklist, 758

    Guinea fowl parasite checklist, 760

    14 Parasites of ungulates, 761

    Deer, 761

    Parasites of the digestive system, 761

    Parasites of the respiratory system, 763

    Parasites of the liver, 766

    Parasites of the circulatory system, 768

    Parasites of the nervous system, 769

    Parasites of the reproductive/urogenital system, 770

    Parasites of the locomotory system, 770

    Parasites of the connective tissue, 771

    Parasites of the integument, 772

    Ectoparasites, 773

    Camels, 773

    Parasites of the digestive system, 773

    Parasites of the respiratory system, 778

    Parasites of the liver, 779

    Parasites of the pancreas, 780

    Parasites of the circulatory system, 780

    Parasites of the nervous system, 784

    Parasites of the locomotory system, 784

    Parasites of the connective tissue, 785

    Ectoparasites, 785

    Llamas, alpacas, guanacos, vicuñas, 787

    Parasites of the digestive system, 787

    Parasites of the respiratory system, 790

    Parasites of the liver, 791

    Parasites of the nervous system, 792

    Parasites of the reproductive/urogenital system, 792

    Parasites of the locomotory system, 792

    Ectoparasites, 792

    Water buffalo, 793

    Parasites of the digestive system, 793

    Parasites of the respiratory system, 796

    Parasites of the liver, 797

    Parasites of the pancreas, 798

    Parasites of the circulatory system, 798

    Parasites of the nervous system, 801

    Parasites of the reproductive/urogenital system, 801

    Parasites of the locomotory system, 801

    Parasites of the connective tissue, 801

    Parasites of the integument, 802

    Ectoparasites, 802

    Deer parasite checklist, 804

    Camel parasite checklist, 808

    Camelid parasite checklist, 811

    Buffalo parasite checklist, 814

    15 Parasites of laboratory animals, 816

    Rabbits, 816

    Parasites of the digestive system, 816

    Parasites of the respiratory system, 826

    Parasites of the liver, 826

    Parasites of the circulatory system, 827

    Parasites of the nervous system, 829

    Parasites of the reproductive/urogenital system, 829

    Parasites of the locomotory system, 829

    Ectoparasites, 830

    Guinea pigs, 833

    Parasites of the digestive system, 833

    Parasites of the respiratory system, 836

    Parasites of the liver, 836

    Parasites of the circulatory system, 836

    Parasites of the nervous system, 836

    Parasites of the reproductive/urogenital system, 836

    Parasites of the locomotory system, 836

    Parasites of the integument, 836

    Ectoparasites, 836

    Rats and mice, 839

    Parasites of the digestive system, 839

    Parasites of the respiratory system, 847

    Parasites of the liver, 847

    Parasites of the circulatory system, 848

    Parasites of the nervous system, 849

    Parasites of the reproductive/urogenital system, 849

    Parasites of the locomotory system, 849

    Ectoparasites, 849

    Primates, 853

    Parasites of the digestive system, 853

    Parasites of the respiratory system, 866

    Parasites of the liver, 868

    Parasites of the pancreas, 869

    Parasites of the circulatory system, 869

    Parasites of the nervous system, 874

    Parasites of the reproductive/urogenital system, 875

    Parasites of the locomotory system, 875

    Parasites of the integument, 876

    Ectoparasites, 876

    Rabbit parasite checklist, 879

    Guinea pig parasite checklist, 881

    Rat parasite checklist, 882

    Mouse parasite checklist, 884

    Primate parasite checklist, 886

    16 Parasites of exotics, 893

    Pigeons, 893

    Parasites of the digestive system, 893

    Parasites of the respiratory system, 897

    Parasites of the circulatory system, 898

    Parasites of the nervous system, 899

    Parasites of the reproductive/urogenital system, 899

    Parasites of the locomotory system, 899

    Parasites of the integument, 899

    Ectoparasites, 900

    Ratites (ostrich, rhea, emu), 903

    Parasites of the digestive system, 903

    Parasites of the respiratory system, 906

    Parasites of the circulatory system, 906

    Parasites of the nervous system, 907

    Ectoparasites, 907

    Reptiles, 908

    Parasites of the digestive system, 908

    Parasites of the respiratory system, 912

    Parasites of the circulatory system, 912

    Parasites of the reproductive/urogenital system, 912

    Ectoparasites, 913

    Pigeon parasite checklist, 915

    Ratite parasite checklist, 918

    17 Facultative ectoparasites and arthropod vectors, 921

    Insects, 921

    Biting and nuisance flies, 928

    Other common hippoboscids, 934

    Myiasis flies, 935

    Ticks, 952

    Hard ticks, 952

    Soft ticks (Argasidae), 968

    Mites, 972

    References and further reading, 975

    Index, 979

  • Mike Taylor

    An independent veterinary consultant, is an honorary Professor in Veterinary Parasitology at the Royal Veterinary College, London, former head of Parasitology at the Central Veterinary Laboratory (VLA), Weybridge and Wildlife Disease Surveillance at Central Science Laboratory York, and Editor-in-Chief of the journal Veterinary Parasitology. He has written more than 250 scientific papers and 20 book chapters in the field.

    Bob Coop

    Is Honorary Fellow of the Moredun Foundation, and former Head of the Division of Parasitology at the Moredun Research Institute, Scotland. He is a former President of the British Association for Veterinary Parasitology and Honorary Fellow of the University of Edinburgh.

    Richard Wall

    Is Professor of Zoology at the University of Bristol, UK. He is a member of the Executive Committee of the World Association for the Advancement of Veterinary Parasitology, and Fellow of the Royal Entomological Society.

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